Funding: State Assignment of the Ministry of Science and Higher Education of the Russian Federation for the Research Centre for Medical Genetics.

Acknowledgements: the authors would like to thank all participants of the expedition survey (donors of samples) and Biobank of North Eurasia for access to DNA collections.

Author contribution: Balanovska EV — management; Agdzhoyan AT — contribution to the expedition survey of Ob Ugrians; Ponomarev GYu — Y-SNP marker genotyping; Ponomarev GYu, Potanina AYu, Adamov DS — statistical and cartographic analysis, manuscript formatting; Balanovska EV, Ponomarev GYu — study design and manuscript writing.

Compliance with ethical standards: the study was approved by the Ethics Committee of the Research Centre for Medical Genetics (protocol No. 1 dated 29 June 2020).